Looking at the $2 \times 2$ and $3 \times 3$ forms of this matrix, we see that:
$\det \begin{bmatrix} t & -a_0 \\ -1 & t-a_1 \end{bmatrix} = t(t-a_1) - a_0 = t^2 - a_1t - a_0$
and, by expansion along the first row:
$\det \begin{bmatrix} t & 0 & -a_0 \\ -1 & t & -a_1 \\ 0 & -1 & t-a_2 \end{bmatrix} = t \times\det \begin{bmatrix} t & -a_1 \\ -1 & t-a_2 \end{bmatrix} + (-a_0) \det\begin{bmatrix} -1 & t \\ 0 & -1 \end{bmatrix}$
$= t[t(t-a_2) - a_1] - a_0 = t^3 - a_2t^2 - a_1t - a_0 $
So it looks like:
$\det \begin{bmatrix} t & 0 & 0 & \cdots & 0 & -a_0 \\ -1 & t & 0 & \cdots & 0 & -a_1 \\ 0 & -1 & t & \cdots & 0 & -a_2 \\ 0 & 0 & -1 & \cdots & 0 & -a_3 \\ \vdots & \vdots & \vdots & \ddots & \vdots & \vdots \\ 0 & 0 & 0 & \cdots & t & -a_{n-1} \\ 0 & 0 & 0 & \cdots & -1 & t-a_n \end{bmatrix} = t^{n+1} - a_nt^n - a_{n-1}t^{n-1} - ... - a_2t^2 - a_1t - a_0$
Which we can prove by induction.
Assume that:
$\det \begin{bmatrix} t & 0 & 0 & \cdots & 0 & -a_0 \\ -1 & t & 0 & \cdots & 0 & -a_1 \\ 0 & -1 & t & \cdots & 0 & -a_2 \\ 0 & 0 & -1 & \cdots & 0 & -a_3 \\ \vdots & \vdots & \vdots & \ddots & \vdots & \vdots \\ 0 & 0 & 0 & \cdots & t & -a_{n-2} \\ 0 & 0 & 0 & \cdots & -1 & t-a_{n-1} \end{bmatrix} = t^{n} - a_{n-1}t^{n-1} - a_{n-2}t^{n-2} - ... - a_2t^2 - a_1t - a_0$
Then, by expansion along the first row:
$\det \begin{bmatrix} t & 0 & 0 & \cdots & 0 & -a_0 \\ -1 & t & 0 & \cdots & 0 & -a_1 \\ 0 & -1 & t & \cdots & 0 & -a_2 \\ 0 & 0 & -1 & \cdots & 0 & -a_3 \\ \vdots & \vdots & \vdots & \ddots & \vdots & \vdots \\ 0 & 0 & 0 & \cdots & t & -a_{n-1} \\ 0 & 0 & 0 & \cdots & -1 & t-a_n \end{bmatrix} = t \det \begin{bmatrix} t & 0 & 0 & \cdots & 0 & -a_1 \\ -1 & t & 0 & \cdots & 0 & -a_2 \\ 0 & -1 & t & \cdots & 0 & -a_3 \\ 0 & 0 & -1 & \cdots & 0 & -a_4 \\ \vdots & \vdots & \vdots & \ddots & \vdots & \vdots \\ 0 & 0 & 0 & \cdots & t & -a_{n-1} \\ 0 & 0 & 0 & \cdots & -1 & t-a_n \end{bmatrix} $
$+ (-1)^{n+1} \times (-a_0)(-1)^n $
$ = t[t^{n} - a_{n}t^{n-1} - a_{n-1}t^{n-2} - ... - a_3t^2 - a_2t - a_1] + (-1)^{2n+1} a_0$
$ = t^{n+1} - a_nt^n - a_{n-1}t^{n-1} - ... - a_2t^2 - a_1t - a_0$
Proof complete.
For part (a), this is just development (Laplace expansion) of the determinant by the first row. Actually the $\det()$ factors should have alternating signs. Since the only occurrences of $x$ are in that first row, all the $\det()$ expressions are constants, and one gets a polynomial of degree at most $n-1$ (from the final term) in $x$.
For part (b), that $P(a_i)=0$ for $i=2,3,\ldots,n$ is just the fact that $P(a_i)$ equals the determinant of the matrix obtained by substituting $a_i$ for $x$, so from the original matrix $a_1$ has been replaced by $a_i$, and as this matrix has its rows $1$ and $i$ identical, its determinant vanishes. all this uses is that the Laplace expansion used commutes with such substitution. Furthermore a polynomial of degree at most $n-1$ with $n-1$ specified roots $a_2,\ldots,a_n$ can only be a scalar multiple of $(x-a_2)\ldots(x-a_n)$.
For part (c), this is just remarking that the $\det()$ in question is $(-1)^{n-1}$ times the determinant of the lower-left $(n-1)\times(n-1)$ submatrix, which determinant precisely matches the definition of $V_{n-1}(a_2,\ldots,a_n)$.
For part (d) write $(-1)^{n-1}\prod_{i=2}^n(x-a_i)=\prod_{i=2}^n(a_i-x)$ to get
$$
V(x,a_2,\ldots,a_n)=
(-1)^{n-1}V_{n-1}(a_2,\ldots,a_n)\prod_{i=2}^n(x-a_i)
=V_{n-1}(a_2,\ldots,a_n)\prod_{i=2}^n(a_i-x),
$$
and then set $x=a_1$ to get
$$
V(a_1,a_2,\ldots,a_n)
=V_{n-1}(a_2,\ldots,a_n)\prod_{i=2}^n(a_i-a_1),
$$
Part (e) applies induction on $n$ to $V_{n-1}(a_2,\ldots,a_n)$ (the starting case is $V_0()=1=\prod_{1\leq i<j\leq 0}1$, an empty product, or if you fear $n=0$ it is $V_1(a)=1=\prod_{1\leq i<j\leq 1}1$, still an empty product), to get
$$
V(a_1,a_2,\ldots,a_n)
=\left(\prod_{2\leq i<j\leq n}(a_j-a_i)\right)\prod_{j=2}^n(a_j-a_1)
=\prod_{1\leq i<j\leq n}(a_j-a_i).
$$
Best Answer
I am very surprised that no solution was given yet.
Denote by $\mathrm{Com}(A)$ the comatrix of $A$. The determinant to be computed is the determinant of the product $D \times \mathrm{Com}(A) \times D$, where $D$ is the diagonal matrix with diagonal entries $((-1)^i)_{1 \le i \le n}$. Since $D^2=I_n$, this matrix is similar to $\mathrm{Com}(A)$, so it as the same determinant as $\mathrm{Com}(A)$.
Since $A \times \mathrm{Com}(A)^\top = (\det A) I_n$, we have $\det(A) \times \det(\mathrm{Com}(A)) = \det((\det A) I_n) = (\det A)^n$.
If $A$ is invertible, we derive $\det(\mathrm{Com}(A)) = (\det A)^{n-1}$.
If $A$ is not invertible and not null, then $\mathrm{Com}(A)$ cannot be invertible since $A \times \mathrm{Com}(A)^\top$ is null, so $\det(\mathrm{Com}(A))$ is null.
If $A$ is null and $n \ge 2$, then $\mathrm{Com}(A)$ is null.
If $A$ is null and $n=1$, then $\mathrm{Com}(A)$ is the $1 \times 1$ matrix with unique entry equal to $1$.
In all cases, we derive $\det(\mathrm{Com}(A)) = (\det A)^{n-1}$.